We acknowledge the herbaria that contributed data to this work: A, AAH, AAS, AAU, ABD, ABH, ABN, ABRN, ABS, ACAD, ACOR, AD, ADW, AFS, AIMS, AJOU, AK, AKPM, AKU, ALCB, ALTA, ALU, AMD, AMDE, AMES, AMNH, AMO, ANA, ANGU, ANSP, AQP, ARAN, ARIZ, ARM, AS, ASDM, ASU, ATCC, AU, AUT, AWH, B, BA, BAA, BAB, BABY, BACP, BAF, BAFC, BAI, BAJ, BAK, BAL, BARC, BAS, BASBG, BBB, BBG, BBS, BC, BCB, BCF, BCN, BCRU, BCW, BDD, BDK, BEL, BEREA, BERN, BEX, BFT, BG, BH, BHCB, BHSC, BIO, BIRA, BIRM, BISH, BKF, BLA, BM, BNM, BNRH, BO, BOCH, BOG, BOL, BOLV, BON, BONN, BOON, BORH, BOTU, BOUM, BOZ, BPI, BR, BREM, BRI, BRIST, BRISTM, BRIT, BRIU, BRLU, BRM, BRNU, BRSN, BRU, BRWK, BRY, BSB, BSIP, BTN, BUF, BUL, BUT, C, CAM, CAMU, CAN, CANB, CAS, CATIE, CAY, CBG, CBM, CBS, CDBI, CEN, CEPEC, CESJ, CGE, CGG, CGMS, CHAM, CHAP, CHAS, CHI, CHL, CHR, CHRB, CHSC, CICY, CIIDIR, CINC, CLE, CLEMS, CLF, CLR, CM, CMC, CMM, CNH, CNPO, CNS, COA, COAH, COFC, COI, COL, COLO, CONC, COR, CORD, CP, CPAP, CPUN, CR, CRAI, CRP, CS, CSLA, CSU, CTES, CTESN, CU, CUVC, CUZ, CVRD, CYN, DAL, DAO, DAV, DBG, DBN, DCR, DEE, DES, DFS, DFSM, DGS, DHM, DLF, DMFS, DMNH, DMU, DNA, DOR, DPU, DR, DS, DSM, DSY, DUE, DUKE, DUSS, E, EA, EAC, EBH, ECON, ECU, EGHB, EIF, EIU, EKY, ELN, EMMA, ENCB, EPM, ER, ERA, ERE, ESA, ETH, EXR, F, FAA, FAU, FAUC, FB, FCO, FCQ, FEN, FH, FHO, FI, FLAS, FLOR, FM, FR, FRS, FRU, FSC, FSU, FTG, FUEL, FULD, FURB, G, GA, GAT, GB, GDA, GENT, GEO, GES, GGO, GH, GI, GJO, GL, GLAM, GLM, GMDRC, GMNHJ, GOET, GOW, GRA, GRM, GSW, GUA, GUAM, GW, GZU, H, HA, HAC, HAL, HAM, HAMAB, HAMU, HAO, HAS, HAST, HASU, HAW, HB, HBG, HBR, HDD, HEID, HGI, HGM, HIB, HIP, HITBC, HIWNT, HLU, HME, HNT, HNUB, HO, HPL, HRCB, HRP, HSS, HSU, HTN, HU, HUA, HUAL, HUEFS, HUEM, HUFU, HUJ, HUSA, HUT, HWB, HXBH, HYO, IAA, IAC, IAN, IB, IBF, IBGE, IBK, IBSC, ICEL, ICESI, ICN, IEA, IFO, ILL, ILLS, INB, INEGI, INM, INPA, INV, IPA, IPRN, ISC, ISE, ISKW, ISL, ISTC, ISU, IZAC, JACA, JBAG, JBGP, JCT, JE, JUA, JYV, K, KAND, KATH, KCS, KGY, KHD, KIEL, KLE, KMN, KMNH, KOELN, KOR, KPABG, KPM, KR, KSP, KSTC, KSU, KTU, KU, KUN, KYO, L, LA, LAE, LAGU, LANC, LAU, LBG, LCN, LCR, LD, LDS, LE, LEA, LEB, LEI, LES, LI, LIL, LINN, LISC, LISE, LISI, LISU, LIV, LIVU, LL, LLN, LMA, LMJ, LMU, LOB, LOJA, LP, LPAG, LPB, LPD, LPS, LSR, LSU, LSUM, LTB, LTN, LTR, LU, LUA, LW, LYJB, LZ, M, MA, MACB, MACF, MAF, MAK, MAL, MAN, MANCH, MARY, MASS, MB, MBA, MBH, MBK, MBM, MBML, MCNS, MEL, MELU, MEN, MERL, MEXU, MFA, MFU, MG, MGC, MHA, MICH, MIL, MIN, MISS, MJG, MMMN, MNA, MNE, MNHM, MNHN, MO, MOR, MPU, MPUC, MSB, MSC, MSE, MSTR, MSUB, MSUN, MT, MTMG, MU, MUB, MUCV, MUR, MVFA, MVFQ, MVJB, MVM, MW, MY, N, NA, NAC, NAS, NCC, NCCBH, NCCE, NCCH, NCE, NCH, NCSC, NCU, NE, NEBC, NH, NHA, NHG, NHM, NHMC, NHT, NM, NMB, NMLU, NMNL, NMR, NMSU, NMW, NOT, NOU, NSPM, NSW, NT, NTS, NU, NUM, NWOSU, NY, O, OBI, OCLA, ODU, OKL, OKLA, OS, OSA, OSC, OSH, OULU, OWU, OXF, P, PACA, PAMP, PAR, PCU, PDD, PE, PEL, PERTH, PEUFR, PFC, PGM, PH, PI, PKDC, PLAT, PLH, PLYP, PMA, PMSP, PNG, PNH, POLL, POM, PORT, PR, PRC, PRE, PRU, PSU, PSY, PTH, PVNH, PY, QCA, QCNE, QFA, QK, QM, QMC, QRS, QUE, R, RAMM, RAS, RB, RBR, RDG, RDS, REG, RELC, RENO, RFA, RIOC, RM, RNG, RSA, RYU, S, SALA, SAM, SAN, SANT, SAPS, SASK, SAV, SBBG, SBT, SCAR, SCFS, SD, SDN, SDSU, SEL, SEV, SF, SFD, SFS, SFV, SGO, SHD, SHIN, SHYB, SI, SIM, SING, SIU, SJC, SJRP, SJSU, SLBI, SLSK, SMDB, SMF, SNM, SNUA, SOM, SP, SPF, SPN, SPSF, SQF, SRFA, SRGH, SSMF, STA, STDCM, STE, STI, STL, STR, STS, STU, SUN, SUU, SUVA, SVG, SWT, SZU, TAA, TAAM, TAES, TAI, TAIF, TALL, TAM, TAMU, TAN, TASH, TBI, TCD, TEF, TENN, TEPB, TEX, TFC, TI, TKPM, TNP, TNS, TO, TOGO, TOR, TOYA, TRA, TRH, TROM, TRT, TRTE, TRTS, TTN, TUB, TUCH, TULS, U, UADY, UAM, UB, UBA, UBC, UBT, UC, UCI, UCMM, UCNW, UCR, UCS, UCSA, UCSB, UCSC, UEA, UEC, UESC, UFG, UFMA, UFMT, UFP, UFRJ, UFRN, UFS, UGDA, UH, UI, UJAT, ULM, ULS, UME, UMO, UNA, UNB, UNCC, UNEX, UNM, UNR, UNSL, UNSW, UPCB, UPEI, UPNA, UPNG, UPP, UPS, US, USAS, USC, USCH, USF, USJ, USM, USNC, USP, USZ, UT, UTC, UTEP, UTLH, UU, UV, UVIC, UWO, V, VAL, VALD, VALPL, VDB, VEN, VIT, VMSL, VPI, VT, W, WAG, WAR, WARMS, WAT, WB, WCR, WCUH, WELT, WFU, WHN, WIES, WII, WIN, WIS, WM, WMNH, WOH, WOLL, WOS, WS, WSCO, WSY, WTU, WU, WXM, YA, YAMA, YK, YRK, Z, ZCM, ZMT, ZSS, ZT.